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Statement - CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952158 - Disclosure - Nature of the Business link:presentationLink link:calculationLink link:definitionLink 9952159 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 9952160 - Disclosure - Marketable Securities link:presentationLink link:calculationLink link:definitionLink 9952161 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 9952162 - Disclosure - Property and Equipment, Net link:presentationLink link:calculationLink link:definitionLink 9952163 - Disclosure - Accrued Expenses and Other Current Liabilities link:presentationLink link:calculationLink link:definitionLink 9952164 - Disclosure - Common Stock and Preferred Stock link:presentationLink link:calculationLink link:definitionLink 9952165 - Disclosure - Stock-Based Compensation link:presentationLink link:calculationLink link:definitionLink 9952166 - Disclosure - Income Taxes link:presentationLink link:calculationLink link:definitionLink 9952167 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 9952168 - Disclosure - Leases link:presentationLink link:calculationLink link:definitionLink 9952169 - Disclosure - Collaboration and License Agreements link:presentationLink link:calculationLink link:definitionLink 9952170 - Disclosure - Employee Benefit Plan link:presentationLink link:calculationLink link:definitionLink 9952171 - Disclosure - Net Income (Loss) per Share link:presentationLink link:calculationLink link:definitionLink 9952172 - Disclosure - Segment Information link:presentationLink link:calculationLink link:definitionLink 9952173 - Disclosure - Subsequent Events link:presentationLink link:calculationLink link:definitionLink 9955511 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9955512 - Disclosure - Summary of Significant Accounting Policies (Tables) link:presentationLink link:calculationLink link:definitionLink 9955513 - Disclosure - Marketable Securities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955514 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955515 - Disclosure - Property and Equipment, Net (Tables) link:presentationLink link:calculationLink link:definitionLink 9955516 - Disclosure - Accrued Expenses and Other Current Liabilities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955517 - Disclosure - Common Stock and Preferred Stock (Tables) link:presentationLink link:calculationLink link:definitionLink 9955518 - Disclosure - Stock-Based Compensation (Tables) link:presentationLink link:calculationLink link:definitionLink 9955519 - Disclosure - Income Taxes (Tables) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - 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Contingencies (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Marketable Securities - Amortized Cost (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Marketable Securities - Amortized Cost (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Marketable Securities - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Fair Value Measurements (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Property and Equipment, Net (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Accrued Expenses and Other Current Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Common Stock and Preferred Stock - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Common Stock and Preferred Stock - Schedule of Shares Reserved for Future Issuance under Equity Compensation Plans (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Stock-Based Compensation - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Stock-Based Compensation - Stock-Based Compensation (Details) link:presentationLink link:calculationLink link:definitionLink 9955537 - Disclosure - Stock-Based Compensation - Stock Option Valuation (Details) link:presentationLink link:calculationLink link:definitionLink 9955538 - Disclosure - Stock-Based Compensation - Stock Option Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955539 - Disclosure - Stock-Based Compensation - Restricted Stock Awards (Details) link:presentationLink link:calculationLink link:definitionLink 9955540 - Disclosure - Income Taxes - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955541 - 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Leases - Maturities of Operating Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955548 - Disclosure - Leases - Maturities of Operating Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955549 - Disclosure - Collaboration and License Agreements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 9955550 - Disclosure - Employee Benefit Plan (Details) link:presentationLink link:calculationLink link:definitionLink 9955551 - Disclosure - Net Income (Loss) per Share - Schedule of Basic and Diluted Net Loss per Share Attributable to Common Stockholders (Details) link:presentationLink link:calculationLink link:definitionLink 9955552 - Disclosure - Net Income (Loss) per Share - Schedule of Antidilutive Securities Excluded from Computation of Net Loss per Share (Details) link:presentationLink link:calculationLink link:definitionLink 9955553 - Disclosure - Segment Information - 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